Subtribe akuatis

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Ito, M., T. Yahara, R. M. King, K. Watanabe, S. Oshita, J. Yokoyama, and D. J. Crawford. 2000. Molecular phylogeny of Eupatorieae (Asteraceae) estimated from cpDNA RFLP and its implication for the polyploid origin hypothesis of the tribe. Journal of Plant Research 113: 91-96.

King, R. M. and H. Robinson. 1987. The genera of the Eupatorieae (Asteraceae). Monographs in Systematic Botany, Missouri Botanical Garden 22: 1-581.

Schilling, E. E., P. B. Cox, and J. L. Panero. 1999. Chloroplast DNA restriction site data support a narrowed interpretation of Eupatorium (Asteraceae). Plant Systematics and Evolution 219: 209-223.

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